Loren D. Coen, Ph.D.


Affiliate Research Professor

Loren D. Coen Ph.D. 


Research Empasis

Marine ecology, benthic ecology, shellfish biology, restoration 

Current and Planned research projects:

  • oyster reef ecology, marine habitat restoration, their functioning and associated ecosystem services
  • remote sensing of intertidal habitats, monitoring of intertidal oyster reefs
  • real-time water quality monitoring
  • benthic ecology, plant-animal interactions, landscape ecology, living shorelines 

Related links:






Selected Publications:

  • Coen, L.D. and A. Humphries, 2017. Oyster-generated marine habitats: their services, enhancement, restoration, and monitoring. In: S. Murphy and S. Allison, Eds., Routledge Handbook of Ecological and Environmental Restoration. Taylor & Francis Group, Routledge, Cambridge, U.K.
  • Coen, L.D., and R.E. Grizzle, 2016.  Bivalve Molluscs. Pp. 89-109.  In: M.J. Kennish, Ed.  Encyclopedia of Estuaries. Springer, Dordrecht, Heidelberg, New York, London.
  • National Research Council (NRC), 2016.  Effective Monitoring to Evaluate Ecological Restoration in the Gulf of Mexico.  Committee on Effective Approaches for Monitoring and Assessing Gulf of Mexico Restoration Activities, National Academies Press, Washington, D.C., 265 pp. 
  • Milbrandt, E.C., M. Thompson, L.D. Coen, R.E. Grizzle, and K. Ward, 2015.  A multiple habitat restoration strategy in a semi-enclosed Florida embayment, combining hydrologic restoration, mangrove propagule plantings and oyster substrate additions.  Ecological Engineering 83:394–404.   doi:10.1016/j.ecoleng.2015.06.043
  • Coen, L.D., and M.J. Bishop.  The ecological role of infectious diseases of molluscs: from populations to ecosystems.  In: S. Ford, N. Carrasco and R. Anderson, Eds., Molluscan diseases and disease processes.  J. of Invertebrate Pathology Special Issue 13:177-211.
  • Baggett, L.P., S.P. Powers, R.D. Brumbaugh, L.D Coen, B. DeAngelis, J. Greene, B. Hancock, S. Morlock, B. Allen, D. Breitburg, D. Bushek, J.H. Grabowski, R. Grizzle, T. Grosholz, M. La Peyre, M. Luckenbach, K. McGraw, M.F. Piehler, S. Westby, and P.S.E. zu Ermgassen, 2015.  Setting guidelines for evaluating performance of oyster habitat restoration.  Restoration Ecology 23:737:745.
  • Coen, L.D., 2014. Biorock™, electricity, and other restoration approaches.  A review of: Innovative Methods of Marine Ecosystem Restoration, Edited by Thomas Goreau and Robert Trench. Ecology 95:790-791.
  • Garcia, R.N., K.W. Chung, P.B. Key, L. Burnett, L.D. Coen, and M.E. DeLorenzo, 2014.  Toxicity of mosquito control insecticides in larval and juvenile Eastern oysters and hard clams, and interactive effects of hypoxia and elevated CO2. Archives of Environmental Contamination and Toxicology 66:450–462.
  • Martin, N., L. Coen, A.J. Martignette, E. Milbrandt, J. Siwicke, and M. Thompson, 2013. Effectiveness of anti-fouling coatings in Southwest Florida’s estuarine and marine waters, with emphasis on real-time observing systems. Florida Sci. 76:259-274.
  • Grizzle, R.E., and L.D. Coen, 2013.  Slow-down and reach out (and we’ll be there): a response to “shellfish as living infrastructure” by Kate Orff.  Ecological Restoration 31:325-329.
  • Baggett, L.P., S.P. Powers, R. Brumbaugh, L.D. Coen, B. DeAngelis, J. Green, B. Hancock, and S. Morlock, 2013.  A handbook for monitoring guidelines and criteria for judging the performance of oyster restoration projects.  TNC Report, Arlington, VA.
  • Milbrandt, E.C., R.D. Bartleson, L.D. Coen, A. Rybak, M.A. Thompson, J. DeAngelo, and P. Stevens, 2012.  Local and regional effects of reopening a tidal inlet on estuarine water quality, seagrass habitat, and fish assemblages.  Continental Shelf Research 41:1–16.
  • Ermgassen, P.S.E. zu, M.D. Spalding, P. Banks, B. Blake, L. Coen, B. Dumbauld, S. Geiger, J.H. Grabowski, R. Grizzle, M. Luckenbach, K. McGraw, B. Rodney, J. Ruesink, S. Powers, and R. Brumbaugh, 2012. Historical ecology with real numbers: Past and present extent and biomass of an imperilled estuarine ecosystem. Proc. Royal Acad. Lond., B. 279 (1742):3393-3400.
  • Coen, L.D., B.R Dumbauld, and M.L. Judge, 2011.  Expanding shellfish aquaculture: a review of the ecological services provided by and impacts of native and cultured bivalves in shellfish-dominated ecosystems.  In: Shellfish aquaculture and the environment, S.E. Shumway, Ed., Wiley-Blackwell
  • Beck, M.W., R.D. Brumbaugh, L. Airoldi, A. Carranza, L.D. Coen, C. Crawford, O. Defeo, G.J. Edgar, B. Hancock., M.C. Kay, H.S. Lenihan, M.W. Luckenbach, C.L. Toropova, G. Zhang, and X. Guo, 2011.  Oyster reefs at risk and recommendations for conservation, restoration and management.  BioScience 61:107–116.
  • Hadley, N.H., M. Hodges, D.H. Wilber, and L.D. Coen, 2010.  Evaluating intertidal oyster reef development in South Carolina using associated faunal indicators.  Restoration Ecology 18:691-701.
  • Beck, M.W., R.D. Brumbaugh, L. Airoldi, A. Carranza, L.D. Coen, C. Crawford, O. Defeo, G.J. Edgar, B. Hancock, M. Kay, H. Lenihan, M.W. Luckenbach, C.L. Toropova, and G. Zhang, 2009.  Shellfish reefs at risk: a global analysis of problems and solutions, The Nature Conservancy, 52pp.
  • Brumbaugh, R.D. and L.D. Coen, 2009. Contemporary approaches for small-scale oyster reef restoration to address substrate versus recruitment limitation: a review and comments relevant for the Olympia oyster, Ostrea lurida (Carpenter, 1864).  J. Shellfish Res. 28:147-161.
  • Brumbaugh, R.D. and L.D. Coen, 2009.  Contemporary approaches for small-scale oyster reef restoration to address substrate versus recruitment limitation: a review and comments relevant for the Olympia oyster, Ostrea lurida (Carpenter, 1864).  J. Shellfish Res. 28:147-161.
  • Coen, L.D., R.D. Brumbaugh, D. Bushek, R. Grizzle, M.W. Luckenbach, M.H. Posey, S.P. Powers, and G. Tolley, 2007.  AS WE SEE IT.  A broader view of ecosystem services related to oyster restoration.  Mar Ecol. Prog. Ser. 341:303–307.
  • Coen, L.D. and R. Grizzle, ASMFC, 2007.  The Importance of Habitat Created by Shellfish and Shell Beds Along the Atlantic Coast of the U.S., with contributions by J. Lowery and K.T. Paynter, Jr., Habitat Management Series #8, 108pp.
  • Bolton-Warberg, M., L.D. Coen and J. Weinstein, 2007.  Acute toxicity and acetylcholinesterase inhibition in grass shrimp (Palaemonetes pugio) and oysters (Crassostrea virginica) exposed to the organophosphate dichlorvos: laboratory and field studies.  Archives of Environmental Contamination and Toxicology 52:207-216.
  • Burrows, F., J.M. Harding, R. Mann, R. Dame and L. Coen, 2005.  Chapter 4, Restoration monitoring of oyster reefs, pages 4.2-4.73.  In: G.W., Thayer, T.A. McTigue, R.J. Salz, D.H. Merkey, F.M. Burrows, and P.F. Gayaldo, (Eds.), Science-Based Restoration Monitoring of Coastal Habitats, Volume Two: Tools for Monitoring Coastal Habitats. NOAA Coastal Ocean Program Decision Analysis Series No. 23. NOAA National Centers for Coastal Ocean Science, Silver Spring, M.D.